Joel works as a smuggler with his friend Tess in a quarantine zone in Boston. In the 20 years that follow, much of civilization is destroyed by the infection survivors live in heavily policed quarantine zones, independent settlements, and nomadic groups. Joel, his brother Tommy, and Sarah attempt to flee the chaos. One night, an outbreak of a mutant Cordyceps fungus ravages the United States, which transforms its human hosts into cannibalistic monsters. In 2013, Joel is a single father living near Austin, Texas with his twelve-year-old daughter Sarah.
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